About the Authors

Junji Otani

Contributed equally to this work with: Junji Otani, Hironobu Kimura

Current address: Graduate School of Biostudies, Kyoto University, Yoshidakonoecho, Sakyo-ku, Kyoto, Japan

Affiliation Graduate School of Engineering, Kyoto University, Katsura, Nishikyo-ku, Kyoto, Japan

Hironobu Kimura

Contributed equally to this work with: Junji Otani, Hironobu Kimura

Affiliation Institute for Protein Research, Osaka University, Suita, Osaka, Japan

Jafar Sharif

Affiliation RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Kanagawa, Japan

Takaho A. Endo

Affiliation RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Kanagawa, Japan

Yuichi Mishima

Affiliation Institute for Protein Research, Osaka University, Suita, Osaka, Japan

Toru Kawakami

Affiliation Institute for Protein Research, Osaka University, Suita, Osaka, Japan

Haruhiko Koseki

Affiliation RIKEN Center for Integrative Medical Sciences, Tsurumi-ku, Yokohama, Kanagawa, Japan

Masahiro Shirakawa

Affiliations Graduate School of Engineering, Kyoto University, Katsura, Nishikyo-ku, Kyoto, Japan, CREST, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Isao Suetake

Affiliations Institute for Protein Research, Osaka University, Suita, Osaka, Japan, CREST, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Shoji Tajima

tajima@protein.osaka-u.ac.jp

Affiliation Institute for Protein Research, Osaka University, Suita, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JO IS H. Kimura ST. Performed the experiments: JO IS H. Kimura YM JS TAE. Analyzed the data: JO H. Kimura IS JS H. Koseki ST TAE. Contributed reagents/materials/analysis tools: TK JO H. Kimura IS YM. Wrote the manuscript: ST IS JO H. Kimura MS H. Koseki.