About the Authors

Garth Mowat

garth.mowat@gov.bc.ca

Affiliation Natural Resource Science Section, Ministry of Forests, Lands and Natural Resource Operations, Nelson, British Columbia, Canada

Douglas C. Heard

Affiliation Fish and Wildlife Section, Ministry of Forests, Lands and Natural Resource Operations, Prince George, British Columbia, Canada

Carl J. Schwarz

Affiliation Department of Statistics and Actuarial Sciences, Simon Fraser University, Burnaby, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GM DCH. Performed the experiments: GM. Analyzed the data: CJS GM. Contributed reagents/materials/analysis tools: GM DCH CJS. Wrote the paper: GM DCH CJS.