About the Authors

Lin Zhang

Contributed equally to this work with: Lin Zhang, Yongxin Fan

Affiliation Department of Life Science and Biotechnology, Nanyang Normal University, Nanyang, China

Yongxin Fan

Contributed equally to this work with: Lin Zhang, Yongxin Fan

Affiliation Department of Life Science and Biotechnology, Nanyang Normal University, Nanyang, China

Haoying Zheng

Affiliation Department of Life Science and Biotechnology, Nanyang Normal University, Nanyang, China

Fengguang Du

Affiliation State Key Laboratory of Motor Bio-fuel technology, Henan Tianguan Group Co. Ltd., Nanyang, China

Ke-qin Zhang

Affiliation Laboratory for Conservation and Utilization of Bio-Resources, and Key Laboratory for Microbial Resources of the Ministry of Education, Yunnan University, Kunming, China

Xiaowei Huang

Affiliation Laboratory for Conservation and Utilization of Bio-Resources, and Key Laboratory for Microbial Resources of the Ministry of Education, Yunnan University, Kunming, China

Linfeng Wang

Affiliation State Key Laboratory of Motor Bio-fuel technology, Henan Tianguan Group Co. Ltd., Nanyang, China

Man Zhang

Affiliation State Key Laboratory of Motor Bio-fuel technology, Henan Tianguan Group Co. Ltd., Nanyang, China

Qiuhong Niu

qiuhongniu@gmail.com

Affiliation Department of Life Science and Biotechnology, Nanyang Normal University, Nanyang, China

Competing Interests

The authors have the following interests: co-authors Fengguang Du, Linfeng Wang and Man Zhang are employed by State Key Laboratory of Motor Bio-fuel technology, Henan Tianguan Group Co. Ltd., Nanyang, China. This study is partly funded by a research grant from State Key Laboratory of Motor Bio-fuel technology (No. 2013014), Henan Tianguan Enterprise Group Co., Ltd., There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: QN LZ. Performed the experiments: YF HZ MZ. Analyzed the data: QN LZ FD LW. Contributed reagents/materials/analysis tools: FD LW. Wrote the paper: QN LZ. Revised the paper: K-qZ XH.