About the Authors

Mickey S. Tivers

Affiliation Department of Veterinary Clinical Sciences, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Ian Handel

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Adam G. Gow

Affiliation Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom

Vicky J. Lipscomb

Affiliation Department of Veterinary Clinical Sciences, Royal Veterinary College, Hatfield, Hertfordshire, United Kingdom

Rajiv Jalan

Affiliation UCL Institute for Liver and Digestive Health, UCL Medical School, London, United Kingdom

Richard J. Mellanby

Richard.Mellanby@ed.ac.uk

Affiliations Royal (Dick) School of Veterinary Studies and The Roslin Institute, The University of Edinburgh, Roslin, Midlothian, United Kingdom, MRC Centre for Inflammation Research, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MST IH AGG RJ VJL RJM. Performed the experiments: MST VJL. Analyzed the data: RJM IH. Contributed reagents/materials/analysis tools: N/A. Wrote the manuscript: RJM RJ MST IH.