About the Authors

Lijuan Zhou

Affiliations Horticultural Research Laboratory, Agricultural Research Service, US Department of Agriculture, Fort Pierce, Florida, United States of America, Indian River Research and Education Center, University of Florida, Fort Pierce, Florida, United States of America

Charles A. Powell

Affiliation Indian River Research and Education Center, University of Florida, Fort Pierce, Florida, United States of America

Wenbin Li

Affiliation Center for Plant Health, Science and Technology, Animal and Plant health Inspection Service, US Department of Agriculture, Beltsville, Maryland, United States of America

Mike Irey

Affiliation Southern Garden Citrus, U. S. Sugar Corp., Clewiston, Florida, United States of America

Yongping Duan

yongping.duan@ars.usda.gov

Affiliation Horticultural Research Laboratory, Agricultural Research Service, US Department of Agriculture, Fort Pierce, Florida, United States of America

Competing Interests

The authors declare there is no Competing Interest although they have a co-author who is employed by a commercial company, Southern Garden Citrus, U. S. Sugar Corp., Clewiston, Florida, United States of America. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LZ YD. Performed the experiments: LZ. Analyzed the data: LZ WL YD. Contributed reagents/materials/analysis tools: CP MI YD. Wrote the paper: LZ WL YD.