About the Authors

Mirjam T. W. van der Sluijs

mirjam.vandersluijs@merck.com

Affiliation Research and Development, MSD Animal Health, Boxmeer, The Netherlands

Dianne P. H. Schroer-Joosten

Affiliation Research and Development, MSD Animal Health, Boxmeer, The Netherlands

Aicha Fid-Fourkour

Affiliation Research and Development, MSD Animal Health, Boxmeer, The Netherlands

Mieke P. Vrijenhoek

Affiliation Animal Services and Pathology, MSD Animal Health, Boxmeer, The Netherlands

Isolde Debyser

Affiliation CoVeToP, Consultancy in Veterinary and Toxicological Pathology, Enghien, Belgium

Véronique Moulin

Affiliation Research and Development, MSD Animal Health, Boxmeer, The Netherlands

Rob J. M. Moormann

Affiliations Central Veterinary Institute, Animal Sciences Group, Wageningen University and Research centre, Lelystad, The Netherlands, Department of Infectious Diseases and Immunology, Virology Division, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands

Abraham J. de Smit

Affiliation Research and Development, MSD Animal Health, Boxmeer, The Netherlands

Competing Interests

This study was funded by MSDAH. M.T.W. van der Sluijs, D.P.H Schroer-Joosten, A. Fid-Fourkour, M.P. Vrijenhoek, V. Moulin and A.J. de Smit are employed Intervet International B.V., which is part of Merck. I. Debeyser is employed by CoVeToP, (Consultancy in Veterinary and Toxicological Pathology). There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: MS RM VM AS. Performed the experiments: MS DS AF MV ID. Analyzed the data: MS MV ID RM AS. Contributed reagents/materials/analysis tools: VM. Wrote the paper: MS RM AS.