About the Authors

Sara E. Gombash

Affiliations Graduate Program in Neuroscience, University of Cincinnati, Cincinnati, Ohio, United States of America, Department of Translational Science and Molecular Medicine, Michigan State University, Grand Rapids, Michigan, United States of America

Fredric P. Manfredsson

Affiliation Department of Translational Science and Molecular Medicine, Michigan State University, Grand Rapids, Michigan, United States of America

Christopher J. Kemp

Affiliation Department of Translational Science and Molecular Medicine, Michigan State University, Grand Rapids, Michigan, United States of America

Nathan C. Kuhn

Affiliation Department of Translational Science and Molecular Medicine, Michigan State University, Grand Rapids, Michigan, United States of America

Sheila M. Fleming

Affiliation Departments of Psychology and Neurology, University of Cincinnati, Cincinnati, Ohio, United States of America

Ann E. Egan

Affiliation Graduate Program in Neuroscience, University of Cincinnati, Cincinnati, Ohio, United States of America

Laura M. Grant

Affiliation Departments of Surgery and Communications Sciences and Disorders, University of Wisconsin, Madison, Wisconsin, United States of America

Michelle R. Ciucci

Affiliation Departments of Surgery and Communications Sciences and Disorders, University of Wisconsin, Madison, Wisconsin, United States of America

Jeffrey P. MacKeigan

Affiliation Laboratory of Systems Biology, Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Caryl E. Sortwell

caryl.sortwell@hc.msu.edu

Affiliation Department of Translational Science and Molecular Medicine, Michigan State University, Grand Rapids, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SEG FPM SMF MRC JPM CES. Performed the experiments: SEG FPM CJK NCK SMF AEE LMG MRC JPM CES. Analyzed the data: SEG CES AEE SMF LMG MRC. Contributed reagents/materials/analysis tools: CES FPM SMF MRC JPM. Wrote the manuscript: SEG FPM SMF AEE LMG MRC JPM CES NCK CJK.