About the Authors

Kohei Sakamoto

Affiliations Laboratory of Veterinary Ethology, The University of Tokyo, Tokyo, Japan, Laboratory of Neurobiology, National Institute of Agrobiological Sciences, Ibaraki, Japan

Yoshihiro Wakabayashi

Affiliation Laboratory of Neurobiology, National Institute of Agrobiological Sciences, Ibaraki, Japan

Takashi Yamamura

Affiliation Laboratory of Neurobiology, National Institute of Agrobiological Sciences, Ibaraki, Japan

Tomomi Tanaka

Affiliation Laboratory of Veterinary Reproduction, Tokyo University of Agriculture and Technology, Tokyo, Japan

Yukari Takeuchi

Affiliation Laboratory of Veterinary Ethology, The University of Tokyo, Tokyo, Japan

Yuji Mori

Affiliation Laboratory of Veterinary Ethology, The University of Tokyo, Tokyo, Japan

Hiroaki Okamura

hokamu@affrc.go.jp

Affiliation Laboratory of Neurobiology, National Institute of Agrobiological Sciences, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HO KS. Performed the experiments: KS YW TY TT. Analyzed the data: KS YW HO. Contributed reagents/materials/analysis tools: KS YW TY. Wrote the manuscript: HO KS. Significant contribution to data interpretation and edition of the manuscript: YT YM.