About the Authors

Zoltan Beck

Affiliations U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America, Henry M. Jackson Foundation for the Advancement of Military Medicine, Silver Spring, Maryland, United States of America

Linda L. Jagodzinski

Affiliation U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Michael A. Eller

Affiliations U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America, Henry M. Jackson Foundation for the Advancement of Military Medicine, Silver Spring, Maryland, United States of America

Doris Thelian

Affiliations U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America, Henry M. Jackson Foundation for the Advancement of Military Medicine, Silver Spring, Maryland, United States of America

Gary R. Matyas

Affiliation U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Anjali N. Kunz

Affiliation U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Carl R. Alving

calving@hivresearch.org

Affiliation U.S. Military HIV Research Program, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZB CRA GRM. Performed the experiments: ZB LLJ MAE DT. Analyzed the data: ZB LLJ MAE DT. Contributed reagents/materials/analysis tools: ZB LLJ MAE DT. Wrote the paper: ZB CRA GRM LLJ MAE. Coordinated the recruitment of the volunteers: ANK.