About the Authors

Joseba M. Garrido

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Patricia Vazquez

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Elena Molina

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Jose M. Plazaola

Affiliation Diputación Foral de Gipuzkoa, Donostia-San Sebastián, Gipuzkoa, Spain

Iker A. Sevilla

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Maria V. Geijo

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Marta Alonso-Hearn

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Ramon A. Juste

rjuste@neiker.net

Affiliation NEIKER-Tecnalia, Animal Health Department, Derio, Bizkaia, Spain

Competing Interests

NEIKER-Tecnalia runs a project for paratuberculosis vaccine improvement funded by the company producing SILIRUM. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: RJ JG. Performed the experiments: JP IS MG PV MA EM. Analyzed the data: RJ. Wrote the paper: JG RJ.