About the Authors

Toshihiro Sakurai

Contributed equally to this work with: Toshihiro Sakurai, So Nakagawa, Min-Su Kim

Current address: Laboratory of Radiophysiology, Department of Life Science, Yokohama College of Pharmacy, Yokohama, Kanagawa, Japan

Affiliation Laboratory of Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

So Nakagawa

Contributed equally to this work with: Toshihiro Sakurai, So Nakagawa, Min-Su Kim

Affiliation Department of Molecular Life Science, Tokai University School of Medicine, Kanagawa, Japan

Min-Su Kim

Contributed equally to this work with: Toshihiro Sakurai, So Nakagawa, Min-Su Kim

Affiliation Laboratory of Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Hanako Bai

Affiliation Laboratory of Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Rulan Bai

Affiliation Laboratory of Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Junyou Li

Affiliation Animal Resource Center, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Kwan-Sik Min

Affiliation Animal Biotechnology, Hankyong National University, Kyonggi-do, Korea

Atsushi Ideta

Affiliation Zen-Noh ET Center, Hokkaido, Japan

Yoshito Aoyagi

Affiliation Zen-Noh ET Center, Hokkaido, Japan

Kazuhiko Imakawa

akaz@mail.ecc.u-tokyo.ac.jp

Affiliation Laboratory of Animal Breeding, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Competing Interests

The authors wish to declare the affiliation of a commercial company Zen-noh ET Center has nothing to do with sharing data or materials in this publication. Thus, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TS M-SK K-SM YA KI. Performed the experiments: TS M-SK HB RB J-YL AI. Analyzed the data: TS SN K-SM HB J-YL YA KI. Contributed reagents/materials/analysis tools: SN AI YA KI. Wrote the paper: SN M-SK RB KI.