About the Authors

Masayuki Ushio

ushima@ecology.kyoto-u.ac.jp

Affiliations Center for Ecological Research, Kyoto University, Otsu, Shiga, Japan, Department of Soil Science, University of Wisconsin-Madison, Madison, Wisconsin, United States of America

Takeshi Miki

Affiliation Institute of Oceanography, National Taiwan University, Taipei, Taiwan

Teri C. Balser

Affiliations Department of Soil Science, University of Wisconsin-Madison, Madison, Wisconsin, United States of America, Department of Soil and Water Science, University of Florida IFAS, Gainesville, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MU TM TCB. Performed the experiments: MU. Analyzed the data: MU TM. Contributed reagents/materials/analysis tools: MU TCB. Wrote the paper: MU TM TCB.