About the Authors

Eliot T. McKinley

Contributed equally to this work with: Eliot T. McKinley, Huiling Liu

Affiliations Vanderbilt University Institute of Imaging Science, Vanderbilt University, Nashville, Tennessee, United States of America, Department of Biomedical Engineering, Vanderbilt University, Nashville, Tennessee, United States of America

Huiling Liu

Contributed equally to this work with: Eliot T. McKinley, Huiling Liu

Affiliation Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

W. Hayes McDonald

Affiliation Department of Biochemistry, Vanderbilt University, Nashville, Tennessee, United States of America

Weifeng Luo

Affiliation Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Ping Zhao

Affiliation Vanderbilt University Institute of Imaging Science, Vanderbilt University, Nashville, Tennessee, United States of America

Robert J. Coffey

Affiliations Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Medicine, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

Steven K. Hanks

Affiliations Department of Cell and Developmental Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America

H. Charles Manning

henry.c.manning@vanderbilt.edu

Affiliations Vanderbilt University Institute of Imaging Science, Vanderbilt University, Nashville, Tennessee, United States of America, Department of Biomedical Engineering, Vanderbilt University, Nashville, Tennessee, United States of America, Vanderbilt Ingram Cancer Center, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Program in Chemical and Physical Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Neurosurgery, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America, Department of Radiology and Radiological Sciences, Vanderbilt University Medical Center, Nashville, Tennessee, United States of America;

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ETM HL RJC SKH HCM. Performed the experiments: ETM HL WHM WL PZ. Analyzed the data: ETM HL SKH HCM. Wrote the manuscript: ETM HL RJC SKH HCM.