About the Authors

Hein Min Tun

Affiliation School of Biological Sciences, The University of Hong Kong, Hong Kong, Hong Kong SAR

Nathalie France Mauroo

Affiliations Department of Pathology, The University of Hong Kong, Hong Kong, Hong Kong SAR, Hong Kong Wildlife Health Foundation, Hong Kong, Hong Kong SAR

Chan San Yuen

Affiliation Clinical Laboratory, Veterinary Center, Ocean Park Corporation, Hong Kong, Hong Kong SAR

John Chi Wang Ho

Affiliation School of Biological Sciences, The University of Hong Kong, Hong Kong, Hong Kong SAR

Mabel Ting Wong

Affiliation School of Biological Sciences, The University of Hong Kong, Hong Kong, Hong Kong SAR

Frederick Chi-Ching Leung

fcleung@hkucc.hku.hk

Affiliations School of Biological Sciences, The University of Hong Kong, Hong Kong, Hong Kong SAR, Bioinformatics Center, Nanjing Agricultural University, Nanjing, China

Competing Interests

Chan San Yuen is an employee of Ocean Park Corporation. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HMT FCL. Performed the experiments: HMT NFM CSY JCWH. Analyzed the data: HMT MTW FCL. Contributed reagents/materials/analysis tools: HMT JCWH MTW FCL. Wrote the paper: HMT NFM CSY JCWH MTW FCL.