About the Authors

Kyo Chul Koo

Affiliation Department of Urology and Urological Science Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Doo Hee Shim

Affiliation Department of Microbiology, Yonsei University College of Medicine, Seoul, Republic of Korea

Chang Mo Yang

Affiliations Department of Microbiology, Yonsei University College of Medicine, Seoul, Republic of Korea, Brain Korea 21 Project for Medical Sciences, Yonsei University College of Medicine, Seoul, Republic of Korea

Saet-Byul Lee

Affiliations Department of Microbiology, Yonsei University College of Medicine, Seoul, Republic of Korea, Brain Korea 21 Project for Medical Sciences, Yonsei University College of Medicine, Seoul, Republic of Korea

Shi Mun Kim

Affiliation ATgen, Sungnam, Republic of Korea

Tae Young Shin

Affiliation Department of Urology and Urological Science Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Kwang Hyun Kim

Affiliation Department of Urology and Urological Science Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Ho Geun Yoon

Affiliations Brain Korea 21 Project for Medical Sciences, Yonsei University College of Medicine, Seoul, Republic of Korea, Department of Biochemistry and Molecular Biology, Severance Medical Research Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Koon Ho Rha

Affiliation Department of Urology and Urological Science Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Jae Myun Lee

sjhong346@yuhs.ac (SJH); jaemyun@yuhs.ac (JML)

Affiliations Department of Microbiology, Yonsei University College of Medicine, Seoul, Republic of Korea, Brain Korea 21 Project for Medical Sciences, Yonsei University College of Medicine, Seoul, Republic of Korea

Sung Joon Hong

sjhong346@yuhs.ac (SJH); jaemyun@yuhs.ac (JML)

Affiliation Department of Urology and Urological Science Institute, Yonsei University College of Medicine, Seoul, Republic of Korea

Competing Interests

An author, SMK, is an employee at ATgen who helped with the experimental process. However, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KCK HGY KHR JML SJH. Performed the experiments: KCK DHS CMY SL SMK. Analyzed the data: KCK TYS KHK. Contributed reagents/materials/analysis tools: SMK HGY KHR JML. Wrote the paper: KCK. Dual corresponding contribution: JML SJH.