About the Authors

Mayo Yasugi

Affiliations Department of Virology, Research Institute for Microbial Diseases (RIMD), Osaka University, Suita, Osaka, Japan, Graduate School of Life and Environmental Sciences, Osaka Prefecture University, Izumisano, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Ritsuko Kubota-Koketsu

Affiliations Department of Virology, Research Institute for Microbial Diseases (RIMD), Osaka University, Suita, Osaka, Japan, Kanonji Institute, The Research Foundation for Microbial Diseases of Osaka University, Kanonji, Kagawa, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Akifumi Yamashita

Affiliation Department of Genome Informatics, RIMD, Osaka University, Suita, Osaka, Japan

Norihito Kawashita

Affiliation Department of Environmental Pharmacometrics, Graduate School of Pharmaceutical Sciences, Osaka University, Suita, Osaka, Japan

Anariwa Du

Affiliations Department of Virology, Research Institute for Microbial Diseases (RIMD), Osaka University, Suita, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Ryo Misaki

Affiliations Applied Microbiology Laboratory, International Center of Biotechnology, Osaka University, Suita, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Motoki Kuhara

Affiliations Ina Laboratory, Medical & Biological Laboratories Corporation, Ltd., Ina, Nagano, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Naphatsawan Boonsathorn

Affiliations Department of Medical Sciences, Ministry of Public Health, Muang, Nonthaburi, Thailand, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Kazuhito Fujiyama

Affiliations Applied Microbiology Laboratory, International Center of Biotechnology, Osaka University, Suita, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Yoshinobu Okuno

Affiliation Kanonji Institute, The Research Foundation for Microbial Diseases of Osaka University, Kanonji, Kagawa, Japan

Takaaki Nakaya

Current address: Department of Infectious Diseases, Kyoto Prefectural University of Medicine, Kyoto, Japan

Affiliations International Research Center for Infectious Diseases, RIMD, Osaka University, Suita, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Kazuyoshi Ikuta

ikuta@biken.osaka-u.ac.jp

Affiliations Department of Virology, Research Institute for Microbial Diseases (RIMD), Osaka University, Suita, Osaka, Japan, The Japan Science and Technology Agency/Japan International Cooperation Agency, Science and Technology Research Partnership for Sustainable Development (JST/JICA, SATREPS), Tokyo, Japan

Competing Interests

The authors have the following interests. Motoki Kuhara is employed by Ina Laboratory, Medical & Biological Laboratories Corporation, Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: MY KI. Performed the experiments: MY RKK AD RM NB. Analyzed the data: MY AY NK KF YO TN KI. Contributed reagents/materials/analysis tools: MK. Wrote the manuscript: MY KI.