About the Authors

Jiansheng Guo

Affiliation Department of Molecular and Cell Biology, School of Life Science and Technology, Tongji University, Shanghai, China

Yongming Yuan

Affiliation Firmenich Company, Shanghai, China

Zhixue Liu

Affiliation Department of Molecular and Cell Biology, School of Life Science and Technology, Tongji University, Shanghai, China

Jian Zhu

zhujian1@tongji.edu.cn

Affiliation Department of Molecular and Cell Biology, School of Life Science and Technology, Tongji University, Shanghai, China

Competing Interests

One of the authors is employed by a commercial company: Firmenich Company. However the article has been allowed for publication and it does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JG YY JZ. Performed the experiments: JG. Analyzed the data: ZL JZ. Contributed reagents/materials/analysis tools: YY JZ. Wrote the paper: JG.