About the Authors

Shigeyuki Fujimoto

Contributed equally to this work with: Shigeyuki Fujimoto, Naoya Itsumura

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Naoya Itsumura

Contributed equally to this work with: Shigeyuki Fujimoto, Naoya Itsumura

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Tokuji Tsuji

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Yasumi Anan

Affiliation Laboratory of Chemical Toxicology and Environmental Health, Showa Pharmaceutical University, Machida, Tokyo, Japan

Natsuko Tsuji

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Yasumitsu Ogra

Affiliations Laboratory of Chemical Toxicology and Environmental Health, Showa Pharmaceutical University, Machida, Tokyo, Japan, High Technology Research Center, Showa Pharmaceutical University, Machida, Tokyo, Japan

Tomoki Kimura

Affiliation Department of Toxicology, Faculty of Pharmaceutical Sciences, Setsunan University, Hirakata, Osaka, Japan

Yusaku Miyamae

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Seiji Masuda

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Masaya Nagao

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Taiho Kambe

kambe1@kais.kyoto-u.ac.jp

Affiliation Division of Integrated Life Science, Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SF NI TT T. Kambe. Performed the experiments: SF NI TT YA NT YO T. Kambe. Analyzed the data: SF NI TT NT T. Kambe. Contributed reagents/materials/analysis tools: T. Kimura YM SM MN T. Kambe. Wrote the paper: SF NI TT T. Kambe.