About the Authors

Hai-Lin Zhang

Contributed equally to this work with: Hai-Lin Zhang, Yu-Zhen Zhang

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Yu-Zhen Zhang

Contributed equally to this work with: Hai-Lin Zhang, Yu-Zhen Zhang

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Wei-Hong Yang

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Yun Feng

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Roger S. Nasci

Affiliation Division of Vector-Borne Diseases, Center for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Jie Yang

Affiliation Dehong Prefecture Center for Disease Control and Prevention, Mangshi, Yunnan, P.R. China

Yong-Hua Liu

Affiliation Ruili City Center for Disease Control and Prevention, Ruili, Yunnan, P.R. China

Chao-Liang Dong

Affiliation Dehong Prefecture Center for Disease Control and Prevention, Mangshi, Yunnan, P.R. China

Shi Li

Affiliation Ruili City Center for Disease Control and Prevention, Ruili, Yunnan, P.R. China

Bao-Sen Zhang

Affiliation Dehong Prefecture Center for Disease Control and Prevention, Mangshi, Yunnan, P.R. China

Zheng-Liu Yin

Affiliation Ruili City Center for Disease Control and Prevention, Ruili, Yunnan, P.R. China

Pi-Yu Wang

Affiliation Yunnan Institute of Parasitic Diseases, Simao, Yunnan, P.R. China

Shi-Hong Fu

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Ming-Hua Li

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Fen Liu

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Juan Zhang

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Jie Sun

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Can-Wei Li

Affiliation Yunnan Institute of Endemic Disease Control and Prevention/ Yunnan Provincial Center of Virus and Rickettsia Research, Dali, Yunnan, P.R. China

Xiao-Yan Gao

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Hong Liu

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Huan-Yu Wang

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Lyle R. Petersen

Affiliation Division of Vector-Borne Diseases, Center for Disease Control and Prevention, Fort Collins, Colorado, United States of America

Guo-Dong Liang

gdliang@hotmail.com

Affiliation State Key Laboratory for Infectious Disease Prevention and Control, Institute for Viral Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, P.R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GDL HLZ RSN LRP HYW. Performed the experiments: HLZ YZZ WHY YF JY YHL CLD SL BSZ ZLY PYW SHF MHL FL JZ JS CWL XYG HL. Analyzed the data: YZZ HLZ RSN GDL. Wrote the manuscript: HLZ RSN YZZ GDL.