About the Authors

Julie L. Dutton

Affiliation Coridon Pty Ltd, Brisbane, Queensland, Australia

Bo Li

Affiliation Coridon Pty Ltd, Brisbane, Queensland, Australia

Wai-Ping Woo

Affiliation Coridon Pty Ltd, Brisbane, Queensland, Australia

Joshua O. Marshak

Affiliation Department of Medicine, University of Washington, Seattle, Washington, United States of America

Yan Xu

Affiliation Coridon Pty Ltd, Brisbane, Queensland, Australia

Meei-li Huang

Affiliations Department of Laboratory Medicine, University of Washington, Seattle, Washington, United States of America, Vaccine and Infectious Diseases Division, Fred Hutchinson Cancer Research Institute, Seattle, Washington, United States of America

Lichun Dong

Affiliation Department of Medicine, University of Washington, Seattle, Washington, United States of America

Ian H. Frazer

Affiliations Coridon Pty Ltd, Brisbane, Queensland, Australia, Diamantina Institute, University of Queensland, Brisbane, Queensland, Australia

David M. Koelle

viralimm@u.washington.edu

Affiliations Department of Medicine, University of Washington, Seattle, Washington, United States of America, Department of Laboratory Medicine, University of Washington, Seattle, Washington, United States of America, Vaccine and Infectious Diseases Division, Fred Hutchinson Cancer Research Institute, Seattle, Washington, United States of America, Department of Global Health, University of Washington, Seattle, Washington, United States of America, Benaroya Research Institute, Seattle, Washington, United States of America

Competing Interests

The authors have the following interests. Partial funding was received from Jagen Pty Ltd and Coridon Pty Limited. JLD, BL, WPW, YX and IHF are employees of, and have share options in Coridon Pty Ltd. IHF is an inventor on the patent US 2011/0287039 A1, “Expression system for modulating an immune response” and WO 2009/049351A1, “Novel compositions and uses therefor” and is a director and shareholder in Coridon Pty Ltd to which these patents have been assigned. JLD is also an inventor on patent US 2011/0287039 A1 and WO 2009/049351 A1. DMK receives consulting fees from Agenus and has received research support from Coridon Pty Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: JLD IHF DMK. Performed the experiments: BL WW JOM YX MH LD. Analyzed the data: JLD MH LD IHF DMK. Wrote the paper: JLD BL WW IHF DMK.