About the Authors

Muqing Zhang

Affiliations Indian River Research and Education Center, IFAS-UF, Fort Pierce, Florida, United States of America, USDA-ARS, US Horticultural Lab, Fort Pierce, Florida, United States of America, State Key Lab for Conservation and Utilization of Subtropical Agro-bioresources, Guangxi University, Nanning, Guangxi, China

Charles A. Powell

Affiliation Indian River Research and Education Center, IFAS-UF, Fort Pierce, Florida, United States of America

Lesley S. Benyon

Affiliation USDA-ARS, US Horticultural Lab, Fort Pierce, Florida, United States of America

Hui Zhou

Affiliation State Key Lab for Conservation and Utilization of Subtropical Agro-bioresources, Guangxi University, Nanning, Guangxi, China

Yongping Duan

yongping.duan@ars.usda.gov

Affiliation USDA-ARS, US Horticultural Lab, Fort Pierce, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CP YD. Analyzed the data: MZ LB HZ CP. Contributed reagents/materials/analysis tools: CP YD. Wrote the paper: MZ LB YD. Carried out the field studies and the DNA extractions: MZ LB. Drafted the manuscript: MZ LB HZ CP.