About the Authors

Randall J. Mitchell

rjm2@uakron.edu

Affiliation Department of Biology, University of Akron, Akron, Ohio, United States of America

William G. Wilson

Affiliation Department of Biology, Duke University, Durham, North Carolina, United States of America

Karsten G. Holmquist

Affiliation Department of Biological Sciences, University of Wisconsin-Milwaukee, Milwaukee, Wisconsin, United States of America

Jeffrey D. Karron

Affiliation Department of Biological Sciences, University of Wisconsin-Milwaukee, Milwaukee, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJM WGW KGH JDK. Performed the experiments: RJM WGW JDK. Analyzed the data: RJM. Contributed reagents/materials/analysis tools: RJM WGW KGH. Wrote the paper: RJM WGW JDK.