About the Authors

David M. Wright

david.m.wright@cantab.net

Current address: Centre for Public Health, Queen’s University Belfast, Belfast, Northern Ireland, United Kingdom

Affiliation School of Biological Sciences, Queen’s University Belfast, Belfast, Northern Ireland, United Kingdom

Adrian R. Allen

Affiliation Veterinary Sciences Division, Bacteriology Branch, Agri-Food and Biosciences Institute, Belfast, Northern Ireland, United Kingdom

Thomas R. Mallon

Affiliation Veterinary Sciences Division, Bacteriology Branch, Agri-Food and Biosciences Institute, Belfast, Northern Ireland, United Kingdom

Stanley W. J. McDowell

Affiliation Veterinary Sciences Division, Bacteriology Branch, Agri-Food and Biosciences Institute, Belfast, Northern Ireland, United Kingdom

Stephen C. Bishop

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

Elizabeth J. Glass

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

Mairead L. Bermingham

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

John A. Woolliams

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

Robin A. Skuce

Affiliations School of Biological Sciences, Queen’s University Belfast, Belfast, Northern Ireland, United Kingdom, Veterinary Sciences Division, Bacteriology Branch, Agri-Food and Biosciences Institute, Belfast, Northern Ireland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARA SWJM RAS JAW EJG SCB. Performed the experiments: TRM ARA RAS SWJM. Analyzed the data: DMW ARA. Wrote the paper: DMW MLB SCB RAS.