About the Authors

Davin M. Henderson

Contributed equally to this work with: Davin M. Henderson, Matteo Manca

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Matteo Manca

Contributed equally to this work with: Davin M. Henderson, Matteo Manca

Affiliations Laboratory of Persistent Viral Diseases, Rocky Mountain Laboratories (RML), National Institute of Allergy and Infectious Disease, Hamilton, Montana, United States of America, Department of Biomedical Sciences, University of Cagliari, Monserrato, Italy

Nicholas J. Haley

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Nathaniel D. Denkers

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Amy V. Nalls

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Candace K. Mathiason

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Byron Caughey

Affiliation Laboratory of Persistent Viral Diseases, Rocky Mountain Laboratories (RML), National Institute of Allergy and Infectious Disease, Hamilton, Montana, United States of America

Edward A. Hoover

Edward.Hoover@colostate.edu

Affiliation Department of Microbiology, Immunology and Pathology, College of Veterinary Medicine and Biomedical Sciences, Colorado State University (CSU), Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DMH MM EAH BC NJH. Performed the experiments: DMH MM NJH ND AVN. Analyzed the data: DMH MM NJH ND AVN. Contributed reagents/materials/analysis tools: CKM EAH BC. Wrote the manuscript: DMH EAH.