About the Authors

Tapani Viitala

Contributed equally to this work with: Tapani Viitala, Niko Granqvist

tapani.viitala@helsinki.fi

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Niko Granqvist

Contributed equally to this work with: Tapani Viitala, Niko Granqvist

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Susanna Hallila

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Manuela Raviña

Affiliation Centre for Drug Research, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Marjo Yliperttula

Affiliation Division of Biopharmaceutics and Pharmacokinetics, Faculty of Pharmacy, University of Helsinki, Helsinki, Finland

Competing Interests

Concerning the competing interest of Niko Granqvist the authors confirm that his part-time employment at Bionavis Ltd as an Application Scientist does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TV NG MR MY. Performed the experiments: SH NG TV. Analyzed the data: TV NG. Contributed reagents/materials/analysis tools: TV NG MY. Wrote the paper: TV NG.