About the Authors

Cheng Hua

Affiliations Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China, College of Chemistry and Life Science, Huanggang Normal University, Huanggang, China, College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, China

Li Linling

Affiliations Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China, College of Chemistry and Life Science, Huanggang Normal University, Huanggang, China

Cheng Shuiyuan

s_y_cheng@sina.com (CS)

Affiliations Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China, College of Chemistry and Life Science, Huanggang Normal University, Huanggang, China, College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, China

Cao Fuliang

s_y_cheng@sina.com (CS)

Affiliation College of Forest Resources and Environment, Nanjing Forestry University, Nanjing, China

Xu Feng

Affiliation Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China

Yuan Honghui

Affiliations Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China, College of Chemistry and Life Science, Huanggang Normal University, Huanggang, China

Wu Conghua

Affiliations Economic Forest Germplasm Improvement and Comprehensive Utilization of Resources of Hubei Key Laboratories, Hubei Huanggang, China, College of Chemistry and Life Science, Huanggang Normal University, Huanggang, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS CH LL. Performed the experiments: CH YH WC. Analyzed the data: CH XF. Contributed reagents/materials/analysis tools: CS CF. Wrote the paper: CS CH.