About the Authors

Syed M. Faisal

Contributed equally to this work with: Syed M. Faisal, Falong Yan

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Falong Yan

Contributed equally to this work with: Syed M. Faisal, Falong Yan

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Tsai-Tzu Chen

Affiliations Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America, Graduate Institute of Agriculture, National Chiayi University, Chiayi, Taiwan

Nicodemus M. Useh

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Shanguang Guo

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Weiwei Yan

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Shih-Jon Wang

Affiliations Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America, Department of Bioscience Technology, Chang Jung Christian University, Tainan, Taiwan

Amy L. Glaser

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Sean P. McDonough

Affiliation Department of Biomedical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Bhupinder Singh

Affiliation Center for Animal Resources & Education, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Yung-Fu Chang

yc42@cornell.edu.

Affiliation Department of Population Medicine and Diagnostic Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Competing Interests

The correspondent author Yung-Fu Chang is a PLOS ONE Editorial board member. This does not alter his adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SMF YFC. Performed the experiments: SMF FY TTC NMU SG WY SJW ALG SPM BS. Analyzed the data: SMF YFC. Wrote the paper: SMF YFC.