About the Authors

Sanna Koskiniemi

Contributed equally to this work with: Sanna Koskiniemi, Henry S. Gibbons

Affiliation Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden

Henry S. Gibbons

Contributed equally to this work with: Sanna Koskiniemi, Henry S. Gibbons

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Linus Sandegren

Affiliation Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden

Naeem Anwar

Affiliation Department of Microbiology, Tumor and Cell Biology, Karolinska Institute, Solna, Sweden

Gary Ouellette

Affiliations US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America, Science Applications International Corporation, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Stacey Broomall

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Mark Karavis

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Paul McGregor

Affiliations US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America, Science Applications International Corporation, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Alvin Liem

Affiliation OptiMetrics, Inc., Abingdon, Maryland, United States of America

Ed Fochler

Affiliation OptiMetrics, Inc., Abingdon, Maryland, United States of America

Lauren McNew

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Carolyn Nicole Rosenzweig

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Mikael Rhen

Affiliation Department of Microbiology, Tumor and Cell Biology, Karolinska Institute, Solna, Sweden

Evan W. Skowronski

Affiliation US Army Edgewood Chemical Biological Center, Aberdeen Proving Ground, Aberdeen, Maryland, United States of America

Dan I. Andersson

Dan.Andersson@imbim.uu.se

Affiliation Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden

Competing Interests

The authors have declared that no competing interests exist. Two of the authors (AL and EF) are employed by a commercial company (Science Applications International Corporation & OptiMetrics, Inc). This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SK HG LS DIA. Performed the experiments: SK NA GO SB MK PM AL EF LM CNR ES. Analyzed the data: SK HG LS NA GO SB MK PM AL EF LM CNR MR ES DIA. Wrote the paper: SK HG LS MR DIA.