About the Authors

Michaël Bekaert

michael.bekaert@stir.ac.uk

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, Scotland, United Kingdom

Natalie R. Lowe

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

Stephen C. Bishop

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

James E. Bron

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, Scotland, United Kingdom

John B. Taggart

Affiliation Institute of Aquaculture, School of Natural Sciences, University of Stirling, Stirling, Scotland, United Kingdom

Ross D. Houston

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MB SCB JEB JBT RDH. Performed the experiments: NRL JEB JBT RDH. Analyzed the data: MB. Wrote the manuscript: MB NRL SCB JEB JBT RDH.