About the Authors

Nakako Izumi Nakajima

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Holly Brunton

Affiliation Genome Damage and Stability Centre, University of Sussex, Brighton, East Sussex, United Kingdom

Ritsuko Watanabe

Affiliation Japan Atomic Energy Agency, Tokai, Japan

Amruta Shrikhande

Affiliation Genome Damage and Stability Centre, University of Sussex, Brighton, East Sussex, United Kingdom

Ryoichi Hirayama

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Naruhiro Matsufuji

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Akira Fujimori

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Takeshi Murakami

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Ryuichi Okayasu

Affiliation Research Center for Charged Particle Therapy and International Open Laboratory, National Institute of Radiological Sciences, Chiba, Japan

Penny Jeggo

p.a.jeggo@sussex.ac.uk (PJ); as303@sussex.ac.uk (AS)

Affiliation Genome Damage and Stability Centre, University of Sussex, Brighton, East Sussex, United Kingdom

Atsushi Shibata

p.a.jeggo@sussex.ac.uk (PJ); as303@sussex.ac.uk (AS)

Current address: Advanced Scientific Research Leaders Development Unit, Gunma University, Maebashi, Japan

Affiliation Genome Damage and Stability Centre, University of Sussex, Brighton, East Sussex, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PJ RO A. Shibata TM. Performed the experiments: NN A. Shibata HB RW A. Shrikhande RH NM AF. Analyzed the data: NN A. Shibata. Contributed reagents/materials/analysis tools: NN A. Shibata PJ RO. Wrote the paper: PJ A. Shibata.