About the Authors

Joel C. Miller

joel.c.miller.research@gmail.com

Affiliation Departments of Mathematics and Biology, Penn State University, University Park, Pennsylvania, United States of America

Erik M. Volz

Affiliation Department of Epidemiology, University of Michigan, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JCM EMV. Performed the experiments: JCM EMV. Analyzed the data: JCM. Wrote the paper: JCM.