About the Authors

Richard Meitern

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Elin Sild

Affiliations Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,, Molecular Ecology and Evolution Lab, Department of Biology, Lund University, Lund, Sweden

Mari-Ann Lind

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Marju Männiste

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Tuul Sepp

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Ulvi Karu

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Peeter Hõrak

horak@ut.ee

Affiliation Department of Zoology, Institute of Ecology and Earth Sciences, Tartu University, Tartu, Estonia,

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PH RM MM. Performed the experiments: RM MM ES M-AL TS UK PH. Analyzed the data: RM MM ES M-AL PH. Contributed reagents/materials/analysis tools: PH. Wrote the paper: RM MM PH.