About the Authors

Cecile M. Krejsa

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Rick D. Holly

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Mark Heipel

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Ken M. Bannink

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Rebecca Johnson

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Richard Roque

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Jane Heffernan

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Julie Hill

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Lay Chin

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Felecia Wagener

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Faith Shiota

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Katherine Henderson

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Pallavur V. Sivakumar

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Hong-Ping Ren

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Fariba Barahmand-pour

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Don Foster

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Chris Clegg

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Wayne Kindsvogel

Affiliation Department of Research, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Rafael Ponce

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Steven D. Hughes

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Kim Waggie

Kim.Waggie@bms.com

Affiliation Department of Pre-clinical Development, ZymoGenetics, Incorporated, a Bristol-Myers Squibb Company, Seattle, Washington, United States of America

Competing Interests

The authors have read the journal’s policy and have the following conflicts: at the time this research was conducted all the authors were employees of ZymoGenetics, Inc., the commercial funder of the research. However, this does not alter the authors’ adherence to all PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CK RH J. Hill PS DF CC WK RP SH. Performed the experiments: MH KB RJ RR J. Heffernan LC FW FS KH HR FB KW. Analyzed the data: CK RH J. Hill PS DF CC WK RP SH MH KB RJ RR J. Heffernan LC FW FS KH HR FB KW. Wrote the paper: CK RH PS WK SH KW.