About the Authors

María Belén Jiménez-Díaz

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Sara Viera

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Javier Ibáñez

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Teresa Mulet

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Noemí Magán-Marchal

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Helen Garuti

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Vanessa Gómez

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Lorena Cortés-Gil

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Antonio Martínez

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Santiago Ferrer

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

María Teresa Fraile

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Félix Calderón

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Esther Fernández

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Leonard D. Shultz

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Didier Leroy

Affiliation Drug Discovery and Technology, Medicines for Malaria Venture, Geneva, Switzerland

David M. Wilson

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

José Francisco García-Bustos

Current address: Department of Microbiology, Monash University, Clayton, Australia

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Francisco Javier Gamo

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Iñigo Angulo-Barturen

inigo.x.angulo@gsk.com

Affiliation Tres Cantos Medicines Development Campus, GlaxoSmithKline, Tres Cantos, Spain

Competing Interests

María Belén Jiménez-Díaz, Francisco Javier Gamo, Sara Viera, Vanessa Gómez, Noemí Magán-Marchal, Helen Garuti, Lorena Cortés-Gil, Antonio Martínez, Santiago Ferrer, María Teresa Fraile, Félix Calderón, Esther Fernández, David M. Wilson, and Iñigo Angulo-Barturen are employees of GlaxoSmithKline. Javier Ibáñez, Teresa Mulet, and José Francisco García-Bustos were employees of GlaxoSmithKline during the conceptual and experimental phase of the work. Didier Leroy is an employee of the Medicines for Malaria Venture. María Belén Jiménez-Díaz, Sara Viera, Vanessa Gómez, Noemí Magán-Marchal, Helen Garuti, Lorena Cortés-Gil Javier Ibáñez, and Teresa Mulet were funded by the Medicines for Malaria Venture under a Miniportfolio Agreement with GlaxoSmithKline, Diseases of the Developing World, Tres Cantos Medicines Development Campus. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: IAB MBJD. Performed the experiments: SV VG NMM HG LCG MBJD. Analyzed the data: IAB MBJD SV. Contributed reagents/materials/analysis tools: MTF SF FC EF LDS. Wrote the paper: IAB. Reviewed and helped write the manuscript: DMW DL FJG FC MBJD SV. Initial analysis and implementation of strategy: IAB FJG JFGB. Approved the paper: MBJD FJG SV JI TM VG NMM HG LCG AM SF MTF FC EF LDS DL DMW JFGB IAB.