About the Authors

Lei Huang

Contributed equally to this work with: Lei Huang, Lingqian Li

Affiliations Cancer Immunology, Inflammation and Tolerance Program, Cancer Center, Georgia Regents University, Augusta, Georgia, United States of America, Department of Radiology, Georgia Regents University, Augusta, Georgia, United States of America

Lingqian Li

Contributed equally to this work with: Lei Huang, Lingqian Li

Affiliation Cancer Immunology, Inflammation and Tolerance Program, Cancer Center, Georgia Regents University, Augusta, Georgia, United States of America

Kim D. Klonowski

Affiliation Department of Cell Biology, College of Arts and Sciences, University of Georgia, Athens, Georgia, United States of America

S. Mark Tompkins

Affiliation Department of Infectious Diseases, College of Veterinary Medicine, University of Georgia Athens, Georgia, United States of America

Ralph A. Tripp

Affiliation Department of Infectious Diseases, College of Veterinary Medicine, University of Georgia Athens, Georgia, United States of America

Andrew L. Mellor

amellor@gru.edu

Affiliations Cancer Immunology, Inflammation and Tolerance Program, Cancer Center, Georgia Regents University, Augusta, Georgia, United States of America, Department of Medicine, Georgia Regents University, Augusta, Georgia, United States of America

Competing Interests

The authors have read the journal's policy and have the following conflict. David Munn and Andrew Mellor serve as members of the Scientific Advisory Board of NewLink Genetics Inc and receive remuneration from this source for this service. They are full-time members of Georgia Regents University and not employees of NewLink Genetics Inc. The service they provide to NewLink Genetics Inc. does not alter the adherence to all the PLOS ONE policies on sharing data and materials as detailed in the guide for authors.

Author Contributions

Conceived and designed the experiments: LH LL KDK SMT RAT ALM. Performed the experiments: LH LL. Analyzed the data: LH LL ALM. Contributed reagents/materials/analysis tools: LH LL KDK MT RAT ALM. Wrote the paper: LH LL ALM.