About the Authors

Madoka Omae

Contributed equally to this work with: Madoka Omae, Norio Takada

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Nara, Japan

Norio Takada

Contributed equally to this work with: Madoka Omae, Norio Takada

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Nara, Japan

Shohei Yamamoto

Affiliation Graduate School of Biological Sciences, Nara Institute of Science and Technology, Nara, Japan

Hiroyuki Nakajima

Affiliation Department of Cell Biology, National Cerebral and Cardiovascular Center Research Institute, Osaka, Japan

Thomas N. Sato

island1005@bs.naist.jp

Affiliations Graduate School of Biological Sciences, Nara Institute of Science and Technology, Nara, Japan, Department of Biomedical Engineering, Cornell University, Ithaca, New York, United States of America, Centenary Institute, Sydney, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MO NT TNS. Performed the experiments: MO NT SY. Analyzed the data: MO NT SY TNS. Contributed reagents/materials/analysis tools: HN. Wrote the paper: MO NT TNS.