About the Authors

Anne Lamontagne

anne.lamontagne@drexelmed.edu (AL); asm54@me.com (AM)

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Ronald E. Long

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Mary Ann Comunale

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Julie Hafner

Affiliation Immunotope Inc., Doylestown, Pennsylvania, United States of America

Lucy Rodemich-Betesh

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Mengjun Wang

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Jorge Marrero

Affiliation Division of Gastroenterology, University of Michigan, Ann Arbor, Michigan, United States of America

Adrian M. Di Bisceglie

Affiliation Department of Internal Medicine and Saint Louis University Liver Center, Saint Louis University School of Medicine, St Louis, Missouri, United States of America

Timothy Block

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Anand Mehta

anne.lamontagne@drexelmed.edu (AL); asm54@me.com (AM)

Affiliation Drexel University College of Medicine, and Department of Microbiology and Immunology and Drexel Institute for Biotechnology and Virology, Doylestown, Pennsylvania, United States of America

Competing Interests

Author Julie Hafner is affiliated to Immunotope Inc. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: AL REL MAC AM. Performed the experiments: AL REL MAC JH LR-B. Analyzed the data: AL REL MAC AM MW. Contributed reagents/materials/analysis tools: JM AMDB TB. Wrote the paper: AM.