About the Authors

Che-Hung Lin

Affiliation Institute of Marine Biology, National Sun Yat-Sen University, Kaohsiung, Taiwan, R.O.C.

Keryea Soong

Affiliation Institute of Marine Biology, National Sun Yat-Sen University, Kaohsiung, Taiwan, R.O.C.

Tung-Yung Fan

tyfan@nmmba.gov.tw

Affiliations National Museum of Marine Biology and Aquarium, Pingtung, Taiwan, R.O.C., Institute of Marine Biodiversity and Evolution, National Dong Hwa University, Pingtung, Taiwan, R.O.C.

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CHL KYS TYF. Performed the experiments: CHL. Analyzed the data: CHL. Contributed reagents/materials/analysis tools: TYF. Wrote the paper: CHL KYS TYF.