About the Authors

Patrick E. Lendrum

lendpatr@isu.edu

Affiliation Department of Biological Sciences, Idaho State University, Pocatello, Idaho, United States of America

Charles R. Anderson Jr

Affiliation Colorado Division of Parks and Wildlife, Grand Junction, Colorado, United States of America

Kevin L. Monteith

Affiliations Department of Biological Sciences, Idaho State University, Pocatello, Idaho, United States of America, Wyoming Cooperative Fish and Wildlife Research Unit, University of Wyoming, Laramie, Wyoming, United States of America

Jonathan A. Jenks

Affiliation Department of Natural Resource Management, South Dakota State University, Brookings, South Dakota, United States of America

R. Terry Bowyer

Affiliation Department of Biological Sciences, Idaho State University, Pocatello, Idaho, United States of America

Competing Interests

The authors acknowledge funding from multiple commercial sources (Williams Production LMT CO., EnCana Corp., Exxon Mobil Production Co., Shell Petroleum and Marathon Oil Corp) and declare that this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: PEL CRA. Performed the experiments: PEL CRA. Analyzed the data: PEL KLM. Contributed reagents/materials/analysis tools: PEL CRA KLM JAJ RTB. Wrote the paper: PEL CRA KLM JAJ RTB.