About the Authors

Noriyuki Shimizu

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Sachiko Chikahisa

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Yuina Nishi

Affiliations Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan, Student Lab, University of Tokushima Faculty of Medicine, Tokushima, Japan

Saki Harada

Affiliations Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan, Student Lab, University of Tokushima Faculty of Medicine, Tokushima, Japan

Yohei Iwaki

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Hiroaki Fujihara

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Kazuyoshi Kitaoka

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Tetsuya Shiuchi

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Hiroyoshi Séi

sei@tokushima-u.ac.jp

Affiliation Department of Integrative Physiology, Institute of Health Biosciences, The University of Tokushima Graduate School, Tokushima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS. Performed the experiments: NS SC YN SH YI HF KK TS. Analyzed the data: NS. Contributed reagents/materials/analysis tools: SC HF KK TS HS. Wrote the paper: HS NS.