About the Authors

Xiaoli Liu

Affiliations Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China, The Graduate School of Chinese Academy of Sciences, Beijing, P. R. China

Huifeng Wu

hfwu@yic.ac.cn

Affiliation Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China

Chenglong Ji

Affiliations Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China, The Graduate School of Chinese Academy of Sciences, Beijing, P. R. China

Lei Wei

Affiliations Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China, The Graduate School of Chinese Academy of Sciences, Beijing, P. R. China

Jianmin Zhao

Affiliation Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China

Junbao Yu

Affiliation Key Laboratory of Coastal Zone Environmental Processes, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Provincial Key Laboratory of Coastal Zone Environmental Processes, YICCAS, Yantai Shandong, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HW. Performed the experiments: XL CJ LW. Analyzed the data: XL HW JZ. Contributed reagents/materials/analysis tools: HW JZ JY. Wrote the paper: XL.