About the Authors

Frank Pasmans

Contributed equally to this work with: Frank Pasmans, Pascale Van Rooij, An Martel, Sebastiano Salvidio

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Pascale Van Rooij

Contributed equally to this work with: Frank Pasmans, Pascale Van Rooij, An Martel, Sebastiano Salvidio

pascale.vanrooij@ugent.be

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Mark Blooi

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Giulia Tessa

Affiliations Dipartimento di Scienze della Vita e Biologia dei Sistemi, Università degli Studi di Torino, Torino, Italy, Zirichiltaggi – Sardinia Wildlife Conservation, Sassari, Italy

Sergé Bogaerts

Affiliation Waalre, The Netherlands

Giuseppe Sotgiu

Affiliation Zirichiltaggi – Sardinia Wildlife Conservation, Sassari, Italy

Trenton W. J. Garner

Affiliation Institute of Zoology, Zoological Society of London, London, United Kingdom

Matthew C. Fisher

Affiliation Department of Infectious Disease Epidemiology, Imperial College, School of Public Health, London, United Kingdom

Benedikt R. Schmidt

Affiliations Institut für Evolutionsbiologie und Umweltwissenschaften, Universität Zürich, Zürich, Switzerland, Koordinationsstelle für Amphibien- und Reptilienschutz in der Schweiz, Neuchâtel, Switzerland

Tonnie Woeltjes

Affiliation Stichting Reptielen, Amfibieën en Vissen Onderzoek Nederland, Nijmegen, The Netherlands

Wouter Beukema

Affiliation Faculty of Geo-Information Science and Earth Observation, University of Twente, Enschede, The Netherlands

Stefano Bovero

Affiliation Zirichiltaggi – Sardinia Wildlife Conservation, Sassari, Italy

Connie Adriaensen

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Fabrizio Oneto

Affiliation Dipartimento di Scienze della Terra, dell’Ambiente e della Vita, Università di Genova, Genova, Italia

Dario Ottonello

Affiliation Dipartimento di Scienze della Terra, dell’Ambiente e della Vita, Università di Genova, Genova, Italia

An Martel

Contributed equally to this work with: Frank Pasmans, Pascale Van Rooij, An Martel, Sebastiano Salvidio

Affiliation Department of Pathology, Bacteriology and Avian Diseases, Faculty of Veterinary Medicine, Ghent University, Merelbeke, Belgium

Sebastiano Salvidio

Contributed equally to this work with: Frank Pasmans, Pascale Van Rooij, An Martel, Sebastiano Salvidio

Affiliation Dipartimento di Scienze della Terra, dell’Ambiente e della Vita, Università di Genova, Genova, Italia

Competing Interests

The authors confirm hereby that co-authors Matthew C. Fisher and Benedikt R. Schmidt are PLOS ONE Editorial Board members. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FP TWJG MCF SS. Performed the experiments: FP PVR MB GT SB GS TW SB CA FO DO AM SS. Analyzed the data: FP PVR MB GT WB BRS AM SS. Wrote the paper: FP PVR TWJG MCF WB BRS AM SS.