About the Authors

Guus M. Bol

Affiliations Departments of Pathology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands, Department of Radiology and Radiological Science, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Venu Raman

Affiliations Departments of Pathology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands, Department of Radiology and Radiological Science, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America, Department of Oncology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Petra van der Groep

Affiliations Departments of Pathology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands, Division of Internal Medicine and Dermatology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands

Jeroen F. Vermeulen

Affiliation Departments of Pathology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands

Arvind H. Patel

Affiliation MRC, University of Glasgow Centre for Virus Research, Glasgow, United Kingdom

Elsken van der Wall

Affiliation Division of Internal Medicine and Dermatology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands

Paul J. van Diest

P.J.vandiest@umcutrecht.nl

Affiliations Departments of Pathology, University Medical Center Utrecht Cancer Center, Utrecht, The Netherlands, Department of Oncology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

Paul van Diest is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GMB VR EW PD. Performed the experiments: GMB PG JFV AHP PD. Analyzed the data: GMB JFV PD. Contributed reagents/materials/analysis tools: AHP PD. Wrote the paper: GMB PD.