About the Authors

Iina Tuominen

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

Leina Al-Rabadi

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

Dimitris Stavrakis

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

Iordanis Karagiannides

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

Charalabos Pothoulakis

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

James M. Bugni

james.bugni@ferring.com

Current address: Ferring Pharmaceuticals Inc., Parsippany, New Jersey, United States of America

Affiliation Division of Digestive Diseases, David Geffen School of Medicine at University of California in Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JMB CP. Performed the experiments: JMB IT LA DS IK. Analyzed the data: JMB IT. Wrote the paper: JMB IT. Contributed to the intellectual content and approved the final version: IT LA DS IK CP JMB.