About the Authors

Toshiyasu Goto

Contributed equally to this work with: Toshiyasu Goto, Atsushi Sato

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Atsushi Sato

Contributed equally to this work with: Toshiyasu Goto, Atsushi Sato

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Masahiro Shimizu

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Shungo Adachi

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Kiyotoshi Satoh

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Shun-ichiro Iemura

Affiliation Biomedicinal Information Research Center, National Institutes of Advanced Industrial Science and Technology, Kohtoh-ku, Tokyo, Japan

Tohru Natsume

Affiliation Biomedicinal Information Research Center, National Institutes of Advanced Industrial Science and Technology, Kohtoh-ku, Tokyo, Japan

Hiroshi Shibuya

shibuya.mcb@mri.tmd.ac.jp

Affiliation Department of Molecular Cell Biology, Medical Research Institute, Tokyo Medical and Dental University, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS . Performed the experiments: TG AS MS SA KS. Analyzed the data: TG AS MS SA KS. Contributed reagents/materials/analysis tools: SI TN. Wrote the paper: HS TG.