About the Authors

Sung Min Bae

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Hee Jung Kim

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Jun Beom Lee

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Jae Bang Choi

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Tae Young Shin

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Hyun Na Koo

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Jae Young Choi

Affiliation Department of Agricultural Biotechnology, College of Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea

Kwang Sik Lee

Affiliation College of Natural Resources and Life Science, Dong-A University, Busan, Republic of Korea

Yeon Ho Je

Affiliations Department of Agricultural Biotechnology, College of Agriculture and Life Science, Seoul National University, Seoul, Republic of Korea, Research Institute for Agriculture and Life Sciences, Seoul National University, Seoul, Republic of Korea

Byung Rae Jin

Affiliation College of Natural Resources and Life Science, Dong-A University, Busan, Republic of Korea

Sung Sik Yoo

Affiliation Choong-Ang Vaccine Laboratory, Daejeon, Republic of Korea

Soo Dong Woo

sdwoo@cbnu.ac.kr

Affiliation Department of Agricultural Biology, College of Agriculture, Life and Environment Sciences, Chungbuk National University, Cheongju, Republic of Korea

Competing Interests

Sung Sik Yoo is employed by Choong-Ang Vaccine Laboratory. There are no patents, products in development or marketed products to declare. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: SDW. Performed the experiments: SMB HJK JBL JBC TYS. Analyzed the data: HNK JYC KSL. Contributed reagents/materials/analysis tools: YHJ BRJ SSY. Wrote the paper: SMB SDW.