About the Authors

Xiaojie Wu

Affiliation College of Marine Science and Engineering, Tianjin University of Science and Technology, Tianjin, China

Aiyou Huang

Affiliations Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, Graduate School, Chinese Academy of Sciences, Beijing, China

Meiling Xu

Affiliation College of Marine Science and Engineering, Tianjin University of Science and Technology, Tianjin, China

Chao Wang

Affiliations Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China, Graduate School, Chinese Academy of Sciences, Beijing, China

Zhaojun Jia

Affiliation College of Marine Science and Engineering, Tianjin University of Science and Technology, Tianjin, China

Guangce Wang

gcwang@ms.qdio.ac.cn

Affiliations College of Marine Science and Engineering, Tianjin University of Science and Technology, Tianjin, China, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Jianfeng Niu

Affiliation Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GCW JFN. Performed the experiments: XJW JFN MLX CW ZJJ. Analyzed the data: XJW AYH JFN. Contributed reagents/materials/analysis tools: JFN. Wrote the paper: XJW JFN GCW.