About the Authors

Jian-Li Zhao

Contributed equally to this work with: Jian-Li Zhao, Lu Zhang

Affiliations Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Kunming, Yunnan, People’s Republic of China, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Menglun, Yunnan, People’s Republic of China, The Graduate University of The Chinese Academy of Sciences, Beijing, People’s Republic of China

Lu Zhang

Contributed equally to this work with: Jian-Li Zhao, Lu Zhang

Affiliations Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Kunming, Yunnan, People’s Republic of China, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Menglun, Yunnan, People’s Republic of China, The Graduate University of The Chinese Academy of Sciences, Beijing, People’s Republic of China

Selvadurai Dayanandan

Affiliation The Graduate University of The Chinese Academy of Sciences, Beijing, People’s Republic of China

Shivaprakash Nagaraju

Affiliation Department of Biology, Concordia University, Montreal, Quebec, Canada

Dong-Mei Liu

Affiliations Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Kunming, Yunnan, People’s Republic of China, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Menglun, Yunnan, People’s Republic of China

Qiao-Ming Li

lqm@xtbg.ac.cn

Affiliations Laboratory of Plant Phylogenetics and Conservation, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Kunming, Yunnan, People’s Republic of China, Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, The Chinese Academy of Sciences, Menglun, Yunnan, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLZ LZ QML. Performed the experiments: JLZ LZ QML. Analyzed the data: JLZ SD SN. Contributed reagents/materials/analysis tools: JLZ LZ DML SD SN. Wrote the paper: JLZ LZ QML SD SN.