About the Authors

Yan-Guo Sun

*sunyanguo1987@126.com (YGS); bkhou@sdu.edu.cn (BKH)

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Bo Wang

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Shang-Hui Jin

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Xiao-Xia Qu

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Yan-Jie Li

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Bing-Kai Hou

*sunyanguo1987@126.com (YGS); bkhou@sdu.edu.cn (BKH)

Affiliation The Key Lab of Plant Cell Engineering and Germplasm Innovation, Education Ministry of China; School of Life Science, Shandong University, Jinan, Shandong, P. R. China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BKH. Performed the experiments: YGS BW SHJ. Analyzed the data: YGS XXQ YJL. Contributed reagents/materials/analysis tools: BW SHJ XXQ. Wrote the paper: BKH YJL YGS.