About the Authors

Hong Shen

Affiliation Department of Anatomy and Cell Biology, Rush University Medical Center, Chicago, Illinois, United States of America

Ricardo E. Perez

Affiliation Department of Anatomy and Cell Biology, Rush University Medical Center, Chicago, Illinois, United States of America

Batzaya Davaadelger

Affiliation Department of Anatomy and Cell Biology, Rush University Medical Center, Chicago, Illinois, United States of America

Carl G. Maki

Carl_Maki@rush.edu

Affiliation Department of Anatomy and Cell Biology, Rush University Medical Center, Chicago, Illinois, United States of America

Competing Interests

The authors have the following interests: Co-author Carl Maki is a PLOS ONE Editorial Board member. Hong Shen was employed by Hoffman-La Roche when the manuscript was originally submitted. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CGM HS. Performed the experiments: HS RP BD. Analyzed the data: CGM HS. Contributed reagents/materials/analysis tools: HS RP. Wrote the paper: CGM.